A very young anther is made up of homogeneous meristematic cells and more or less oblong in shape. 1. Parietal cells lie immediately below the epidermis. Hypodermal definition: of the hypodermis | Meaning, pronunciation, translations and examples These cells act as sporangial initials. The number of stamens comprised by the androecium is sometimes the same as the number of petals, but often the stamens are more numerous or fewer in number than the petals. These divide by periclinal walls to form inner primary sporogenous cells or archesporium and outer primary parietal cells. In maize anther primordia, the outer (L1) layer produces the epidermis . a. Epidermis: It is single layered and protective in function. The hypodermal cells then secrete a soft and pliant new cuticle, extending new microvilli into it as it is constructed. Anthers typically contain four somatic cell layers encircling the sporogenous cells within each anther locule (Goldberg et al., 1993). A group of hypodermal cells are present at each corner of the young anther. The cells undergo repeated anticlinal divisions to cope up with the rapidly enlarging internal tissues. Other articles where Anther is discussed: angiosperm: The androecium: …terminal saclike structures (microsporangia) called anthers. We took advantage of the jam-1::gfp marker ( Mohler et al., 1998 ), which localizes to the adherens junctions surrounding the seam cells. It soon becomes four lobed. hypodermal: (hī′pə-dûr′məl) adj. Subsequent to the differentiation of parietal cells and sporogenous cells promoted by SPL/NZZ and BAM1/2, a number of genes are involved in the specialization of tapetal cells, including several leucine-rich repeat RLK genes, Lying below the epidermis. Of or relating to the hypodermis. In each of the four lobes, some of the hypodermal cells begin to act as archesporial initials. hypodermal wall cell layers are replaced by microsporocyte-like cells (Hord et al., 2006). Anther wall : The mature anther wall consists of the following layers a. Epidermis b. Endothecium c. Middle layers d. Tapetum. derm n. 1. Gradually the anther becomes a four-lobed structure, in each lobe hypodermal cells become differentiated by their shape and dense contents with distinct nucleus, these cells are called archespon'um or archesporial cells. hypodermic: [ hi″po-der´mik ] 1. beneath the skin; injected into subcutaneous tissues. 2. a subcutaneous, injection; a syringe for making such an injection. The inner (L2) hypodermal cells divide periclinally to produce archesporial cells internally and primary parietal cells externally. Immediately after ecdysis, entry of water into the individual through the gills or gut causes inflation of the body that expands the new cuticle to its final size ( Capen, 1972 , Mykles, 1980 , Taylor and Kier, 2003 ). 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