Anthers typically contain four somatic cell layers encircling the sporogenous cells within each anther locule (Goldberg et al., 1993). 2. derm n. 1. b. (a) Formation of sporangial wall There are generally two pairs of spore-containing sacs … It soon becomes four lobed. hypodermal: (hī′pə-dûr′məl) adj. These cells act as sporangial initials. The hypodermal cells then secrete a soft and pliant new cuticle, extending new microvilli into it as it is constructed. a. Epidermis: It is single layered and protective in function. In maize anther primordia, the outer (L1) layer produces the epidermis . Lying below the epidermis. These divide by periclinal walls to form inner primary sporogenous cells or archesporium and outer primary parietal cells. The inner (L2) hypodermal cells divide periclinally to produce archesporial cells internally and primary parietal cells externally. hypodermic: [ hi″po-der´mik ] 1. beneath the skin; injected into subcutaneous tissues. An epidermal layer of cells that secretes an overlying chitinous cuticle, as in arthropods. A group of hypodermal cells are present at each corner of the young anther. Anther wall : The mature anther wall consists of the following layers a. Epidermis b. Endothecium c. Middle layers d. Tapetum. Of or relating to the hypodermis. Gradually the anther becomes a four-lobed structure, in each lobe hypodermal cells become differentiated by their shape and dense contents with distinct nucleus, these cells are called archespon'um or archesporial cells. Hypodermal definition: of the hypodermis | Meaning, pronunciation, translations and examples hypodermal wall cell layers are replaced by microsporocyte-like cells (Hord et al., 2006). Parietal cells lie immediately below the epidermis. The cells undergo repeated anticlinal divisions to cope up with the rapidly enlarging internal tissues. Immediately after ecdysis, entry of water into the individual through the gills or gut causes inflation of the body that expands the new cuticle to its final size ( Capen, 1972 , Mykles, 1980 , Taylor and Kier, 2003 ). The number of stamens comprised by the androecium is sometimes the same as the number of petals, but often the stamens are more numerous or fewer in number than the petals. 1. In each of the four lobes, some of the hypodermal cells begin to act as archesporial initials. 2. a subcutaneous, injection; a syringe for making such an injection. The seam cells are lateral hypodermal blast cells that donate daughter nuclei to the hypodermal syncytium after each cell division, and eventually fuse in adults (Sulston and Horvitz, 1977). Other articles where Anther is discussed: angiosperm: The androecium: …terminal saclike structures (microsporangia) called anthers. 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